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The slotted knuckle is the basic explanation.  However, a point that is often missed in the historical overview is that there was a considerable amount of incompatibility between link and pin couplers. There were many different designs and many required a variety of specialized connecting links and pins in order to mate.    This led to crews hoarding the loose link and pin hardware so as to have the proper components to mate couplers.  A crew setting out twenty cars might strip all of the links and pins, and add them to their stockpile in the caboose.  The crew arriving to pick up the cars, would have to go through their stockpile to find all the necessary links and pins to install on the couplers of the twenty cars.  Undoubtedly, this fundamental awkward incompatibility problem continued into the changeover phase to automatic couplers as the well-established working practice of the pure link and pin era.
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