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Plywood does move somewhat with humidity changes; although not nearly as much as solid wood.  A 9' run might move as much as a 1/4' over dry to very humid conditions.  Add to this the temperature expansion/contraction of metal rail; and you could have a combination that would cause the problem.  I suspect that the glue you used is really the root of the problem.  Regular white glue seems to work better and the caulk will be even better yet.  Cut a couple of gaps in the straight sections and re glue with regular white clue or caulk.
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